Souls
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Souls are permanent, team-wide buffs that are gained from Soul/Biome Bosses.
You can see which two (2) Souls are active in each match by looking at the map.
It's possible for matches to have only one Soul, or even none.
Soul Bosses spawn at the start of the 2nd and 4th days.
The Soul must be picked up where the boss dies, leaving behind a Biome Collapse (expanding purple zone that scales with the current zone damage).